Fact, Fable, and Darwin, Part 1

The American Enterprise Institute. ^ | February 2005 | By Rodney Stark

[restornu]

I write as neither a creationist nor a Darwinist, but as one who knows what is probably the most disreputable scientific secret of the past century: There is no plausible scientific theory of the origin of species! Darwin himself was not sure he had produced one, and for many decades every competent evolutionary biologist has known that he did not. Although the experts have kept quiet when true believers have sworn in court and before legislative bodies that Darwin's theory is proven beyond any possible doubt, that's not what reputable biologists, including committed Darwinians, have been saying to one another.

Without question, Charles Darwin would be among the most prominent biologists in history even if he hadn't written The Origin of Species in 1859. But he would not have been deified in the campaign to "enlighten" humanity. The battle over evolution is not an example of how heroic scientists have withstood the relentless persecution of religious fanatics. Rather, from the very start it primarily has been an attack on religion by militant atheists who wrap themselves in the mantle of science.

When a thoroughly ideological Darwinist like Richard Dawkins claims, "The theory is about as much in doubt as that the earth goes round the sun," he does not state a fact, but merely aims to discredit a priori anyone who dares to express reservations about evolution. Indeed, Dawkins has written, "It is absolutely safe to say that, if you meet somebody who claims not to believe in evolution, that person is ignorant, stupid, or insane ...."

That is precisely how "Darwin's Bulldog," Thomas Huxley, hoped intellectuals would react when he first adopted the tactic of claiming that the only choice is between Darwin and Bible literalism. However, just as one can doubt Max Weber's Protestant Ethic thesis without thereby declaring for Marxism, so too one may note the serious shortcomings of neo-Darwinism without opting for any rival theory. Modern physics provides a model of how science benefits from being willing to live with open questions rather than embracing obviously flawed conjectures.

What is most clear to me is that the Darwinian Crusade does not prove some basic incompatibility between religion and science. But the even more immediate reality is that Darwin's theory falls noticeably short of explaining the origin of species. Dawkins knows the many serious problems that beset a purely materialistic evolutionary theory, but asserts that no one except true believers in evolution can be allowed into the discussion, which also must be held in secret. Thus he chastises Niles Eldridge and Stephen Jay Gould, two distinguished fellow Darwinians, for giving "spurious aid and comfort to modern creationists."

Dawkins believes that, regardless of his or her good intentions, "If a reputable scholar breathes so much as a hint of criticism of some detail of Darwinian theory, that fact is seized upon and blown up out of proportion." While acknowledging that "the extreme rarity of transitional forms in the fossil record" is a major embarrassment for Darwinism, Stephen Jay Gould confided that this has been held as a "trade secret of paleontology" and acknowledged that the evolutionary diagrams "that adorn our textbooks" are based on "inference ... not the evidence of fossils."

According to Steven Stanley, another distinguished evolutionist, doubts raised by the fossil record were "suppressed" for years. Stanley noted that this too was a tactic begun by Huxley, always careful not to reveal his own serious misgivings in public. Paleontologist Niles Eldridge and his colleagues have said that the history of life demonstrates gradual transformations of species, "all the while really knowing that it does not." This is not how science is conducted; it is how ideological crusades are run.

By Darwin's day it had long been recognized that the fossil evidence showed that there had been a progression in the biological complexity of organisms over an immense period of time. In the oldest strata, only simple organisms are observed. In more recent strata, more complex organisms appear. The biological world is now classified into a set of nested categories. Within each genus (mammals, reptiles, etc.) are species (dogs, horses, elephants, etc.) and within each species are many specific varieties, or breeds (Great Dane, Poodle, Beagle, etc.).

It was well-known that selective breeding can create variations within species. But the boundaries between species are distinct and firm – one species does not simply trail off into another by degrees. As Darwin acknowledged, breeding experiments reveal clear limits to selective breeding beyond which no additional changes can be produced. For example, dogs can be bred to be only so big and no bigger, let alone be selectively bred until they are cats. Hence, the question of where species come from was the real challenge and, despite the title of his famous book and more than a century of hoopla and celebration, Darwin essentially left it unanswered.

After many years spent searching for an adequate explanation of the origin of species, in the end Darwin fell back on natural selection, claiming that it could create new creatures too, if given immense periods of time. That is, organisms respond to their environmental circumstances by slowly changing (evolving) in the direction of traits beneficial to survival until, eventually, they are sufficiently changed to constitute a new species. Hence, new species originate very slowly, one tiny change after another, and eventually this can result in lemurs changing to humans via many intervening species.

Darwin fully recognized that a major weakness of this account of the origin of species involved what he and others referred to as the principle of "gradualism in nature." The fossil record was utterly inconsistent with gradualism. As Darwin acknowledged: "...why, if species have descended from other species by fine gradations, do we not everywhere see innumerable transitional forms? Why is not all nature in confusion instead of the species being, as we see them, well defined?"

Two Solutions

Darwin offered two solutions. Transitional types are quickly replaced and hence would mainly only be observable in the fossil record. As for the lack of transitional types among the fossils, that was, Darwin admitted, "the most obvious and serious objection which can be urged against the theory."

Darwin dealt with this problem by blaming "the extreme imperfection of the geological record." "Only a small portion of the surface of the earth has been geologically explored, and no part with sufficient care." But, just wait, Darwin promised, the missing transitions will be found in the expected proportion when more research has been done. Thus began an intensive search for what the popular press soon called the "missing links."

Today, the fossil record is enormous compared to what it was in Darwin's day, but the facts are unchanged. The links are still missing; species appear suddenly and then remain relatively unchanged. As Steven Stanley reported: "The known fossil record...offers no evidence that the gradualistic model can be valid."

Indeed, the evidence has grown even more contrary since Darwin's day. "Many of the discontinuities [in the fossil record] tend to be more and more emphasized with increased collecting," noted the former curator of historical geology at the American Museum of Natural History. The history of most fossil species includes two features particularly inconsistent with gradualism, Stephen Jay Gould has acknowledged. The first problem is stasis. Most species exhibit no directional change during their tenure on earth. They appear in the fossil record looking much the same as when they disappear. The second problem is sudden appearance. Species do not arise gradually by the steady transformation of ancestors, they appear "fully formed."

These are precisely the objections raised by many biologists and geologists in Darwin's time – it was not merely that Darwin's claim that species arise through eons of natural selection was offered without supporting evidence, but that the available evidence was overwhelmingly contrary. Unfortunately, rather than concluding that a theory of the origin of species was yet to be accomplished, many scientists urged that Darwin's claims must be embraced, no matter what.

In keeping with Darwin's views, evolutionists have often explained new species as the result of the accumulation of tiny, favorable random mutations over an immense span of time. But this answer is inconsistent with the fossil record wherein creatures appear "full-blown and raring to go." Consequently, for most of the past century, biologists and geneticists have tried to discover how a huge number of favorable mutations can occur at one time so that a new species would appear without intermediate types.

However, as the eminent and committed Darwinist Ernst Mayr explained, “The occurrence of genetic monstrosities by mutation ... is well substantiated, but they are such evident freaks that these monsters can only be designated as 'hopeless.' They are so utterly unbalanced that they would not have the slightest chance of escaping elimination through selection. Giving a thrush the wings of a falcon does not make it a better flyer.... To believe that such a drastic mutation would produce a viable new type, capable of occupying a new adaptive zone, is equivalent to believing in miracles.”

The word miracle crops up again and again in mathematical assessments of the possibility that even very simple biochemical chains, let alone living organisms, can mutate into being by a process of random trial and error. For generations, Darwinians have regaled their students with the story of the monkey and the typewriter, noting that given an infinite period of time, the monkey sooner or later is bound to produce Macbeth purely by chance, the moral being that infinite time can perform miracles.

However, the monkey of random evolution does not have infinite time. The progression from simple to complex life forms on earth took place within a quite limited time. Moreover, when competent mathematicians considered the matter, they quickly calculated that even if the monkey's task were reduced to coming up with only a few lines of Macbeth, let alone Shakespeare's entire play, the probability is far, far beyond mathematical possibility. The odds of creating even the simplest organism at random are even more remote – Fred Hoyle and Chandra Wickramasinghe, celebrated cosmologists, calculated the odds as one in ten to the 40,000th power. (Consider that all atoms in the known universe are estimated to number no more than ten to the 80th power.) In this sense, then, Darwinian theory does rest on truly miraculous assumptions.

Perhaps the most amazing aspect of the current situation is that while Darwin is treated as a secular saint in the popular media and the theory of evolution is regarded as the invincible challenge to all religious claims, it is taken for granted among the leading biological scientists that the origin of species has yet to be explained. Writing in Nature in 1999, Eörs Szathmay summarizes that, "The origin of species has long fascinated biologists. Although Darwin's major work bears it as a title, it does not provide a solution to the problem." When Julian Huxley claimed that "Darwin's theory is...no longer a theory but a fact," he surely knew better. But, just like his grandfather, Thomas Huxley, he knew that his lie served the greater good of "enlightenment

[Doctor Stochastic]

Could have been worse; the author could have been the basketball coach.

[longshadow]

Thus outing himself as a Creationist. No other group has to explain their position at the beginning of an article.

Someone writing an article like this and prefacing it with a declaration that they are NOT a Creationist is bit like an essay on the appealing physiques of young boys written by Michael Jackson starting out with a declaration "I am not a pedophile, but...."

[Ichneumon]

But, just wait, Darwin promised, the missing transitions will be found in the expected proportion when more research has been done. Thus began an intensive search for what the popular press soon called the "missing links." Today, the fossil record is enormous compared to what it was in Darwin's day, but the facts are unchanged. The links are still missing; species appear suddenly and then remain relatively unchanged. As Steven Stanley reported: "The known fossil record...offers no evidence that the gradualistic model can be valid."

Um, gee, really? Is the author really that grossly ignorant about the actual state of the fossil record, or is he just lying his face off about it? With anti-evolutionists, it's so hard to tell...

For example:

Example 2: reptile-mammals

Figure 1.4.1. The jaws of three vertebrates—mammal, therapsid, and pelycosaur. A side view of three idealized skulls of mammals, therapsids (mammal-like reptiles), and pelycosaurs (early reptiles). The figure shows the differences between mammal and reptilian jaws and ear-bone structures. The jaw joint is shown as a large black dot, the quadrate (mammalian anvil or incus) is in turquoise, the articular (mammalian hammer or malleus) is in yellow, and the angular (mammalian tympanic annulus) is in pink. Note how, in the reptile, the jaw joint is formed between the blue quadrate and the yellow articular (with the pink angular close by), and how, in the mammal, the jaw joint is formed between the squamosal above and the dentary below. In the reptile, the squamosal is just above and contacting the quadrate. Advanced therapsids have two jaw joints: a reptile-like joint and a mammal-like joint (Figure based on Kardong 2002, pp. 275, reproduced with permission from the publisher, Copyright © 2002 McGraw-Hill)

We also have an exquisitely complete series of fossils for the reptile-mammal intermediates, ranging from the pelycosauria, therapsida, cynodonta, up to primitive mammalia (Carroll 1988, pp. 392-396; Futuyma 1998, pp. 146-151; Gould 1990; Kardong 2002, pp. 255-275). As mentioned above, the standard phylogenetic tree indicates that mammals gradually evolved from a reptile-like ancestor, and that transitional species must have existed which were morphologically intermediate between reptiles and mammals—even though none are found living today. However, there are significant morphological differences between modern reptiles and modern mammals. Bones, of course, are what fossilize most readily, and that is where we look for transitional species from the past. Osteologically, two major striking differences exist between reptiles and mammals: (1) reptiles have at least four bones in the lower jaw (e.g. the dentary, articular, angular, surangular, and coronoid), while mammals have only one (the dentary), and (2) reptiles have only one middle ear bone (the stapes), while mammals have three (the hammer, anvil, and stapes) (see Figure 1.4.1).

Early in the 20^th century, developmental biologists discovered something that further complicates the picture. In the reptilian fetus, two developing bones from the head eventually form two bones in the reptilian lower jaw, the quadrate and the articular (see the Pelycosaur in Figure 1.4.1). Surprisingly, the corresponding developing bones in the mammalian fetus eventually form the anvil and hammer of the unique mammalian middle ear (also known more formally as the incus and malleus, respectively; see Figure 1.4.2) (Gilbert 1997, pp. 894-896). These facts strongly indicated that the hammer and anvil had evolved from these reptilian jawbones—that is, if common descent was in fact true. This result was so striking, and the required intermediates so outlandish, that many anatomists had extreme trouble imagining how transitional forms bridging these morphologies could have existed while retaining function. Young-earth creationist Duane Gish stated the problem this way:

"All mammals, living or fossil, have a single bone, the dentary, on each side of the lower jaw, and all mammals, living or fossil, have three auditory ossicles or ear bones, the malleus, incus and stapes. ... Every reptile, living or fossil, however, has at least four bones in the lower jaw and only one auditory ossicle, the stapes. ... There are no transitional fossil forms showing, for instance, three or two jawbones, or two ear bones. No one has explained yet, for that matter, how the transitional form would have managed to chew while his jaw was being unhinged and rearticulated, or how he would hear while dragging two of his jaw bones up into his ear." (Gish 1978, p. 80)

Figure 1.4.2. A comparison of the ears of reptiles and mammals. The reptile ear is shown on the left, the mammal ear on the right. As in Figure 1.4.1, the quadrate (mammalian anvil or incus) is in turquoise and the articular (mammalian hammer or malleus) is in yellow. The stapes is shown in brown. Note how the relative arrangement of these bones is similar in both taxa, in the order of inner ear-stapes-quadrate-articular.

Gish was incorrect in stating that there were no transitional fossil forms, and he has been corrected on this gaff numerous times since he wrote these words. However, Gish's statements nicely delineate the morphological conundrum at hand. Let's review the required evolutionary conclusion. During their evolution, two mammalian middle ear bones (the hammer and anvil, aka malleus and incus) were derived from two reptilian jawbones. Thus there was a major evolutionary transition in which several reptilian jawbones (the quadrate, articular, and angular) were extensively reduced and modified gradually to form the modern mammalian middle ear. At the same time, the dentary bone, a part of the reptilian jaw, was expanded to form the major mammalian lower jawbone. During the course of this change, the bones that form the hinge joint of the jaw changed identity. Importantly, the reptilian jaw joint is formed at the intersection of the quadrate and articular whereas the mammalian jaw joint is formed at the intersection of the squamosal and dentary (see Figure 1.4.1).

How could hearing and jaw articulation be preserved during this transition? As clearly shown from the many transitional fossils that have been found (see Figure 1.4.3), the bones that transfer sound in the reptilian and mammalian ear were in contact with each other throughout the evolution of this transition. In reptiles, the stapes contacts the quadrate, which in turn contacts the articular. In mammals, the stapes contacts the incus, which in turn contacts the malleus (see Figure 1.4.2). Since the quadrate evolved into the incus, and the articular evolved into the malleus, these three bones were in constant contact during this impressive evolutionary change. Furthermore, a functional jaw joint was maintained by redundancy—several of the intermediate fossils have both a reptilian jaw joint (from the quadrate and articular) and a mammalian jaw joint (from the dentary and squamosal). Several late cynodonts and Morganucodon clearly have a double-jointed jaw. In this way, the reptilian-style jaw joint was freed to evolve a new specialized function in the middle ear. It is worthy of note that some modern species of snakes have a double-jointed jaw involving different bones, so such a mechanical arrangement is certainly possible and functional.

Since Figure 1.4.3 was made, several important intermediate fossils have been discovered that fit between Morganucodon and the earliest mammals. These new discoveries include a complete skull of Hadrocodium wui (Luo et al. 2001) and cranial and jaw material from Repenomamus and Gobiconodon (Wang et al. 2001). These new fossil finds clarify exactly when and how the malleus, incus, and angular completely detached from the lower jaw and became solely auditory ear ossicles.

Recall that Gish stated: "There are no transitional fossil forms showing, for instance, three or two jawbones, or two ear bones" (Gish 1978, p. 80). Gish simply does not understand how gradual transitions happen (something he should understand, obviously, if he intends to criticize evolutionary theory). These fossil intermediates illustrate why Gish's statement is a gross mischaracterization of how a transitional form should look. In several of the known intermediates, the bones have overlapping functions, and one bone can be called both an ear bone and a jaw bone; these bones serve two functions. Thus, there is no reason to expect transitional forms with intermediate numbers of jaw bones or ear bones. For example, in Morganucodon, the quadrate (anvil) and the articular (hammer) serve as mammalian-style ear bones and reptilian jaw bones simultaneously. In fact, even in modern reptiles the quadrate and articular serve to transmit sound to the stapes and the inner ear (see Figure 1.4.2). The relevant transition, then, is a process where the ear bones, initially located in the lower jaw, become specialized in function by eventually detaching from the lower jaw and moving closer to the inner ear.

Figure 1.4.3. A comparison of the jawbones and ear-bones of several transitional forms in the evolution of mammals. Approximate stratigraphic ranges of the various taxa are indicated at the far left (more recent on top). The left column of jawbones shows the view of the left jawbone from the inside of the mouth. The right column is the view of the right jawbone from the right side (outside of the skull). As in Figure 1.4.1, the quadrate (mammalian anvil or incus) is in turquoise, the articular (mammalian hammer or malleus) is in yellow, and the angular (mammalian tympanic annulus) is in pink. For clarity, the teeth are not shown, and the squamosal upper jawbone is omitted (it replaces the quadrate in the mammalian jaw joint, and forms part of the jaw joint in advanced cynodonts and Morganucodon). Q = quadrate, Ar = articular, An = angular, I = incus (anvil), Ma = malleus (hammer), Ty = tympanic annulus, D = dentary. (Reproduced from Kardong 2002, pp. 274, with permission from the publisher, Copyright © 2002 McGraw-Hill)

The above is from 29+ Evidences for Macroevolution, which compiles several hundred transitional fossils, which is itself just a *SMALL* sampling of the ENORMOUS numbers of fine transitional sequences found in the fossil record and well known to anyone who has bothered to CRACK OPEN A BOOK -- or even do a websearch -- in the past 25 years or so... So what's the anti-evolutionists' excuse for remaining abysmally ignorant of such things, and repeatedly making the false claim that there are "no" transitional fossils, etc.?

Here's another look:

Mammal-Like Reptiles

As previously stated, a succession of transitional fossils exists that link reptiles (Class Reptilia) and mammals (Class Mammalia). These particular reptiles are classifie as Subclass Synapsida. Presently, this is the best example of th e transformation of one major higher taxon into another. The morphologic changes that took place are well documented by fossils, beginning with animals essentially 100% reptilian and resulting in animals essentially 100% mammalian. Therefore, I have chosen this as the example to summarize in more detail (Table 1, Fig. 1).

Skulls and jaws of synapsid reptiles and mammals; left column side view of skull; center column top view of skull; right column side view of lower jaw. Hylonomus modified from Carroll (1964, Figs. 2,6; 1968, Figs. 10-2, 10-5; note that Hylonomus is a protorothyrod, not a synapsid). Archaeothyris modified from Reisz (1972, Fig. 2). Haptodus modified from Currie (1977, Figs, 1a, 1b; 1979, Figs. 5a, 5b). Sphenacodo n modified from Romer & Price (1940, Fig. 4f), Allin (1975, p. 3, Fig. 16);note: Dimetrodon substituted for top view; modified from Romer & Price, 1940, pl. 10. Biarmosuchus modified from Ivakhnenko et al. (1997, pl. 65, Figs. 1a, 1B, 2); Alin & Hopson (1992; Fig. 28.4c); Sigogneau & Tchudinov (1972, Figs. 1, 15). Eoarctops modified from Broom (1932, Fig. 35a); Boonstra (1969, Fig. 18). Pristerognathus modified from Broom (1932, Figs 17a, b,c); Boonstra (1963, Fig. 5d). Procynosuchus modified from Allin & Hopson (1992, Fig. 28.4e); Hopson (1987, Fig. 5c); Brink (1963, Fig. 10a); Kemp (1979, Fig. 1); Allin (1975, p. 3, Fig. 14). Thrinaxodon modified from Allin & Hopson (1992, Fig. 28.4f);Parrington (1946, Fig. 1); Allin (1975, p. 3, Fig. 13). Probainognathus modified from Allin & Hopson (1992, Fig. 28.4g); Romer (1970, Fig. 1); Allin (1975, p. 3, Fig. 12). Morga nucodon modified from Kermack, Mussett, & Rigney (1981, Figs. 95, 99a; 1973, Fig. 7a); Allin (1975, p. 3, Fig. 11). Asioryctes modified from Carroll (1988, Fig. 20-3b). Abbreviations: ag = angular; ar = articular; cp = coronoid process; d = dentary; f = lateral temporal fenestra; j = jugal; mm = attachment site for mammalian jaw muscles; o = eye socket; po = post orbital; q = quadrate; rl = reflected lamina; sq = squamosal; ty = tympanic.

TAXONOMY	LATERAL TEMPORAL FENESTRA	LOWER JAW DENTARY	TEETH	LOWER JAW: POST- DENTARY BONES	MIDDLE EAR & JAW ARTICULATION
M: Early Placental mammals Asioryctes Upper Cretaceous	Merged with eye socket; cheek arch bowed out laterally	100% of jaw length is the den- tary; condylar process in contact with squamosal	Fully differentiated teeth; incisors, canines, premolars; one tooth replacement	No post-dentary bones	3 middle ear bones (stapes, incus, malleus) + tympanic; squamosal-dentary jaw joint
L: "Pantothere" mammals Amphitherium Middle/Upper Jurassic	X	100% of jaw length is the den-  tary; condylar process contacts squamosal	Fully differentiated teeth; incisors, canines, premolars; one tooth replacement	Post-dentary bones migrated to middle ear	Probably 3 middle ear bones (stapes, incus, malleus) + tympanic; squamosal-dentary jaw joint
K: Morganucodontid mammals Morganucodon  Upper Triassic & Lower Jurassic	Merged with eye socket; cheeck arch bowed out laterally	100% of jaw length is the den- tary; condylar process expanded posteriorly to make contact with squamosal	Fully differentiated teeth; incisors, canines, premolars; one tooth replacement	20% of jaw length; reflected lamina decreased to narrow ribbon-like horseshoe	Stapes extends from inner ear capsule to quadrate; quadrate tiny; both quadrate-articular and squamosal-dentary jaw joints
J: Chiniquodontid cynodonts Probainognathus Middle Triassic	Much larger than eye socket; 40- 45% of skull length; expanded posterioirly, medially, & laterally; midline of skull narrow sagittal crest; chek arch bowed out laterally	95% of jaw length is the dentary; large coronoid process expanded posteriorly; condylar process expanded posteriorly	Large single canine; cheek teeth multicusped; tooth replacement reduced	20% of jaw length; angular notch widened ventrally; width of main part of angular decreased; reflec - ted lamina decreased to narrow ribbon-like horseshoe	Stapes extends from inner ear capsule to quadrate; quadrate tiny; quadrate-articular joint
I:Galesaurid cynodonts Thrinaxodon Lower Triassic	Much larger than eye socket; 40% of skull length; expanded pos- terioirly, medially, & laterally; midline of skull narrow sagittal crest; chek arch bowed out laterally	85% of jaw length is the dentary; large coronoid process expanded to top of eye socket and pos- teriorly; jaw muscles attached to most of coronoid process	Large single canine; cheek teeth multicusped; tooth replacement reduced	25% of jaw length; angular notch widened ventrally; width of reflec- ted lamina decreased; width of main part of angular decreased	Stapes extends from inner ear capsule to quadrate; quadrate small; quadrate-articular jaw joint
H: Procynosuchid cynodonts Procynosuchus upper Upper Permian	Much larger than eye socket; 40% of skull length; expanded pos- terioirly, medially, & laterally; midline of skull narrow sagittal crest; chek arch bowed out laterally	75-80% of jaw length is the den- tary; coronoid process expanded to near top of eye socket and posteriorly; jaw muscles  attached to dorsal part of coronoid process	Large single canine; cheek teeth multicusped	30% of jaw length; angular notch widened ventrally; width of reflected lamina decreased	Stapes extends from inner ear capsule to quadrate; quadrate small; quadrate-articular jaw joint
G: Early Therocephalians Pristerognathus lower Upper Permian	Larger than eye socket; expanded posteriorly and medially; 30% of skull length	75-80% of jaw length is the den- tary; posterior end of dentary expanded posteriorly and dorsally into narrow blade-like coronoid process; rises to middle of eye socket	Large single canine; other teeth simple cones.	35% of jaw length; angular notch deepened into a cleft; reflected lamina large, broad, blade-like	Stapes extends from inner ear capsule to quadrate; quadrate small; quadrate-articular jaw joint
F: Early Gorgonopsians Eoarctops lower Upper Permian	Slightly larger than eye socket; expanded posteriorly and medially (minimal); 20-25% of skull length	65-75% of jaw length is the den- tary; posterior end of dentary slightly expanded posteriorly and dorsally as incipient coronoid process	Large single canine; other teeth simple cones.	40% of jaw length; angular notch deepened into a cleft; reflected lamina large, broad, blade-like	Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint
E: Eotitanosuchians Sphenacodon Lower Permian	Small; slightly smaller than eye socket; slightly expanded posteriorly and medially	65-75% of jaw length is the den- tary; posterodorsal edge rises broadly but slightly above tooth row	Large single canine; other teeth simple cones.	40% of jaw length; angular notch deepened into a cleft; reflected lamina large, broad, blade-like	Stapes extends from inner ear capsule to quadrate;  quadrate- articular jaw joint
D: Late sphenacodonts Sphenacodon Upper Pennsylvanian	Small; smaller than eye socket; confined to one side of skull	65% of jaw length is the dentary; posterodorsal edge rises broadly but slightly above the tooth row	Enlarged incipient canines; other teeth simple cones	60% of jaw length; venntral edge of angular notched ("angular notch") offsetting a short pro- tusion (reflected lamina)	Stapes extends from inner ear capsule to quadrate; quadrate large and plate-like; quadrate- articular jaw joint
C: Early spenacodonts Haptodus Upper Pennsylvanian	Tiny; smaller than eye socket; confined to one side of skull	65-75% of jaw length is the den- tary; posterodorsal edge rises broadly but slightly above tooth row	Undifferentiated; slightly enlarged incipient canines just behind nares	70% of jaw length; ventral edge of angular with shallow indentation	Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint
B: Early ophiacodonts Archaothyris upper Middle Pennsylvanian	Tiny; smaller than eye socket; confined to one side of skull	x	Undifferentiated; slightly enlarged incipient canines just behind nares	x	Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint
A: Protorothyrids Hylonomus lower Middle Pennsylvanian	Absent	65-75% of jaw length is the den- tary; posterodorsal edge rises broadly but slightly above tooth row	Undifferentiated; slightly enlarged incipient canines just behind nares	70% of jaw length; ventral edge of angular continuous	Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint

Table 1: Morphology of synapsid reptiles and mammals (Note that Hylonomus is a protothyrid, not a synapsid). Data from references cited in text.

Modern reptiles and mammals are very distinctive, easily diagnosable, and do not intergrade. Reptiles are covered by scales, mammals by hair; reptiles are cold-blooded, mammals warm-blooded; reptiles do not suckle their young, mammals have mammary glands; reptiles have sprawling posture, mammals have upright posture. Most of these features are soft part anatomy or physiology that very rarely fossilize (although dinosaur skin impressions are known from Cretaceous sediments, and imprints of mammal hair are known from Eocene bats from Germany; Franzen, 1990). In the fossil record, we must look to skeletal features.

There are many skeletal features which allow us to distinguish the reptiles from the mammals (Carroll, 1988; Table 1, rows A, M). The single most important defining characteristic is the nature of the articulation of the lower jaw to the skull (Simpson, 1959). In reptiles, multiple bones comprise the lower jaw. A small bone at the posterior end of the lower jaw, the articular, articulates with the quadrate bone of the skull (Simpson, 1959; Carroll, 1988). In mammals, one large bone, the dentary, comprises the lower jaw. It articulates with the squamosal bone of the skull (Simpson, 1959; Carroll, 1988).

From comparative anatomy studies, it is certain that most of the bones of the reptiles and mammals are homologous (Crompton & Parker, 1978; Carroll, 1988). Of greatest importance, the middle ear bones of mammals (stapes, incus, malleus, and tympanic) are homologous with several of the skull and jaw bones of reptiles (stapes, quadrate, articular, and angular, respectively; Romer, 1956, p. 33-38, 1970a; Allin, 1975, 1986; Allin & Hopson, 1992; Crompton & Parker, 1978; Hopso n, 1987, 1994; Carroll, 1988). One group of reptiles, the synapsids (Subclass Synapsida), share with the mammals an additional homologous structure: the lateral temporal fenestra, which is an opening in the skull behind the eye socket at the triple junction between the squamosal, jugal , and post orbital bones (Broom, 1932; Frazetta, 1968; Kemp, 1982; Carroll, 1988). A band of bone composed of the jugal and the squamosal is adjacent to the lateral temporal fenestra (Broom, 1932; Kemp, 1982; Carroll, 1988). This is the cheek arch so characteristic of mammal skulls (Broom, 1932; Kemp, 1982; Carroll, 1988). Therefore, synapsids are commonly named the “mammal-like reptiles.”

The presence of diagnosable morphologic differences between reptiles (including the oldest reptiles and the oldest synapsids) and mammals distinguishes them as distinct taxa. This allows us to test evolution by looking for transitional forms between the two. Because many of the bones are homologous, we should find evidence illustrating how these bones were modified over time to become the new bones. Furthermore, these morphologic changes should happen in parallel and in geochronologic succession.

Synapsid reptiles inhabited Pangea from the Middle Pennsylvanian through the Early Jurassic (Kemp, 1982, 1985; Sloan, 1983; Carroll, 1988; Hopson, 1969, 1987, 1994; Hopson & Crompton, 1969; Hotton, et al., 1986; Crompton & Jenkins, 1973; Sidor & Hopson, 1998; Romer & Price, 1940; Broom, 1932; Boonstra, 1963, 1969, 1971; Tchudinov, 1983; Olson, 1944; Tatarinov, 1974; Vyushkov, 1955; Efremov, 1954). From the Early Permian through the Early Triassic, they were the largest and most abundant land animals (Sloan, 1983; Colbert, 1965). Though much less well known to the general public than dinosaurs, one of the “cereal box dinosaurs,” Dimetrodon (the sail-backed reptile), is a synapsid, not a dinosaur (Romer & Price, 1940; Carroll, 1988). The oldest mammals are Late Triassic (Kemp, 1982; Carroll, 1988). Below is a discussion of the geochronologic succession linking synapsids and mammals. The oldest reptiles (named protorothyrids; Carroll, 1964, 1988, p. 192-199) are from the lower Middle Pennsylvanian, and the oldest synapsids (Reisz, 1972) are from the upper Middle Pennsylvanian, both of Nova Scotia. Upper Pennsylvanian and Lower Permian forms are known primarily from the midcontinent and Permian Basin region of the United States (Romer & Price, 1940; Currie, 1977, 1979; Kemp, 1982; Sloan, 1983). The basal Upper Permian forms are known from Russia (Tchudinov, 1960, 1983; Efremov, 1954; Olson, 1962; Sigogneau & Tchudinov, 1972; Ivakhnenko et al., 1997). Most of the Upper Permian and Lower Triassic succession is known from southern Africa, especially the Great Karoo of South Africa (Broom, 1932; Boonstra, 1963, 1969, 1971; Hopson & Kitching, 1972; Kemp, 1982; Sloan, 1983). The Middle Triassic forms are from South America (Romer, 1969a, 1969b, 1970b, 1973; Romer & Lewis, 1973; Bonaparte & Barbarena, 1975), and the Upper Triassic and Lower Jurassic mammals are known from Eurasia (Kermack, Mussett, & Rigney, 1973, 1981; Kemp, 1982). Subsequent Mesozoic mammals are known from all over the world (Simpson, 1928; Lillegraven et al., 1979).

When placed in proper geochronologic succession, the synapsids naturally form a succession of taxa (genera and families) that progressively become more mammal-like and less reptile-like (Kemp, 1982, 1985; Sloan, 1983; Sidor & Hopson, 1998; Hopson, 1987, 1994). Morphologic changes, summarized in Table 1 and Figure 1, affect the entire skeletal anatomy of these animals, but are most clearly displayed in their skulls.

The lateral temporal fenestra increased in size from a tiny opening smaller than the eye socket to a giant opening occupying nearly half the length of the skull. Ultimately, it merged with the eye socket, thus producing the full development of the cheek arch so characteristic of mammals (Broom, 1932; Frazetta, 1968; Kemp, 1982; Sloan, 1983; Hopson, 1987, 1994; Carroll, 1988).

Successively, the relative proportion of the lower jaw comprised of the dentary bone (teeth-bearing bone) gradually increased until the entire lower jaw consisted of the dentary (Kemp, 1982; Sloan, 1983; Carroll, 1988; Hopson, 1987, 1994). In Pennsylvanian and Lower and basal Upper Permian synapsids, the postero-dorsal edge of the lower jaw rose broadly but only slightly above the level of the tooth row (Romer & Price, 1940; Currie, 1977, 1979; Ivakhnenko et al., 1997; Tchudinov, 1960, 1983; Efremov, 1954; Olson, 1962; Sigogneau & Tchudinov, 1972; Hopson, 1987, 1994). In succeeding forms, the posterior part of the dentary expanded dorsally and posteriorly as a blade-like process, and progressively became larger (Broom, 1932; Boonstra, 1963, 1969, 1971; Sigogneau, 1970; Brink, 1963; Kemp, 1979; Hopson, 1987, 1994), forming the coronoid process (Parrington, 1946; Fourie, 1974; Romer, 1969b, 1970b, 1973; Hopson, 1987, 1994) to which the mammalian-type jaw musculature is attached (Barghusen, 1968; Bramble, 1978; Crompton, 1972; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988). Concomitantly, the post-dentary bones progressively reduced in size (Allin, 1975; Crompton, 1972; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988; Hopson, 1987, 1994).

Beginning with the Upper Pennsylvanian sphenacodonts, a notch developed in the angular bone that offsets a projection, the reflected lamina (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Romer & Price, 1940; Currie, 1977, 1979; Kemp, 1982; Sloan, 1983; Carroll, 1988). The reflected lamina first became a large blade-like flange (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Ivakhnenko et al., 1997; Tchudinov, 1960, 1983; Efremov, 1954; Olson, 1962; Sigogneau & Tchudinov, 1972; Broom, 1932; Sigogneau, 1970; Boonstra, 1963, 1969, 1971), and then was progressively reduced to a delicate horseshoe-shaped bone (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Brink, 1963; Parrington, 1946; Fourie, 1974; Romer, 1969b, 1970b, 1973; Kermack, Mussett, & Rigney, 1973, 1981; Kemp, 1979, 1982; Sloan, 1983; Carroll, 1988).

Simultaneously, the quadrate progressively decreased in size (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Kemp, 1982; Sloan, 1983; Carroll, 1988). The articular did not decrease in size much, being small initially, but developed a downward-pointing prong (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Kemp, 1982; Sloan, 1983; Carroll, 1988). In the synapsids, the lower jaw was hinged to the skull by the articular and quadrate bones (Crompton, 1972; Crompton & Parker, 1978; Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994). Thus they are classified as reptiles (Simpson, 1959; Kemp, 1982; Sloan, 1983; Carroll, 1988). As the quadrate and articular became smaller, they were relieved of their solid suture to the dentary and skull (Crompton, 1972; Allin, 1975, 1986; Allin & Hopson, 1992; Hopson, 1987, 1994; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988). A projection of the dentary extended posteriorly and made contact with the squamosal. Morganucodon possessed the mammalian dentary-squamosal jaw joint adjacent to the reptilian articular-quadrate jaw joint (Kermack, Mussett, & Rigney, 1973, 1981; Carroll, 1988). It is classified as the first mammal, but it is a perfect intermediate. Now that a new jaw joint was established, the quadrate and articular were subsequently relieved of that function (Crompton, 1972; Allin, 1975, 1986; Allin & Hopson, 1992; Hopson, 1987, 1994; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988). Ultimately, in Middle and Upper Jurassic mammals, the tiny quadrate, articular, and ring-like angular migrated as a unit to the middle ear where they joined the stapes and became the incus, malleus, and tympanic bones (Allin, 197 5, 1986; Allin & Hopson, 1992; Hopson, 1987, 1994; Kemp, 1982; Sloan, 1983; Carroll, 1988).

Progressively, the teeth became differentiated. The large canines developed first, followed by the development of multicusped cheek teeth, reduced tooth replacement (Osborn & Crompton, 1973; Crompton & Parker, 1978), and finally full y differentiated incisors, canines, premolars, and molars with one tooth replacement during life (Kemp, 1982; Hopson, 1994).

Many other morphologic changes are documented in the fossil record. These demonstrate the morphologic and geochronologic succession from sprawling limb posture to upright limb posture of mammals (Jenkins, 1971; Romer & Lewis, 197 3; Kemp, 1982; Carroll, 1988; Hopson, 1994). As Jenkins (1971, p. 210) stated, “In details of morphology and function, the cynodont post-cranial skeleton should be regarded as neither ‘reptilian’ nor ‘mammalian’ but as transitional between the two classes .” Other changes have been adequately summarized elsewhere (Kemp, 1982; Sloan, 1983; Carroll, 1988; Hopson, 1994). Obviously, fundamental physiologic changes must have taken place as well, many of which are not directly preserved in the fossil record, though some can be inferred from the skeletal anatomy (Findlay, 1968; Kemp, 1982; Sloan, 1983, Carroll, 1988; Hopson, 1994).

This is well documented in the fossil record by a massive volume of incontrovertible data that cannot be explained away. Such large-scale, progressive, continuous, gradual, and geochronologically successive morphologic change (Sidor & Hopson, 1998) is descent with modification, and provides compelling evidence for evolution on a grand scale.

(The above is from The Fossil Record: Evolution or "Scientific Creation", which is yet ANOTHER source the anti-evolutionists are obviously completely ignorant of -- not that that stops them from spouting off falsehoods about the subject anyway...

It never ceases to amaze me how much the anti-evolutionsts *don't* know about the actual evidence. Aren't you supposed to actually *know* something about a topic before you attempt to critique?

[Alamo-Girl]

Thanks for the ping!

[longshadow]

Did you notice those quotes? Those are all common creationist canards that you usually see on every creationist website that purports to have quotes from established scientists on what they think of evolution. I'd bet the Quote Mine Project has each of them already there.

Being that the author is a sociologist, you don't suppose he's doing this on purpose as a reverse "Sokal-type" sting operation to make conservative think-tanks look idiotic? The obvious use of "QuoteMine" type material suggests that he made no effort to hide the shoddiness of his scholarship.

[Doctor Stochastic]

Or like a "seminar caller" to one of the radio talk shows. Each show seems to have its cadre of those. The use the same jargon and make the same (usually fallacious) argument.

[Youngblood]

There is no plausible scientific theory of the origin of species! Darwin himself was not sure he had produced one, and for many decades every competent evolutionary biologist has known that he did not. Although the experts have kept quiet when true believers have sworn in court and before legislative bodies that Darwin's theory is proven beyond any possible doubt, that's not what reputable biologists, including committed Darwinians, have been saying to one another.

Who are these biologists who have allegedly been conspiring for decades behind our backs to keep the charade that is evolutionary biology going?!!

[Youngblood]

What is anti-Christian and/or anti-American about evolutionary biology?

[Ichneumon]

[ Evolutionary Theory ] is still, as it was in Darwin's time, a highly speculative hypothesis entirely without direct factual support

ROFL!!! Only if you close your eyes real tight and pretend not to see the *mountains* of evidence which provides overwhelming support for evolutionary biology... Where do the anti-evolutionists *get* these folks?

So, what's going on here?

An anti-evolutionist is misrepresenting the state of the science, that's what's going on.

He still can't explain HOW life came about.

Claiming to read minds of the dead now? Fascinating.

If Ernst Mayr is the "end all" of neo-Darwinism, how can his version of Darwin's theory be so weak as to not withstand basic scrutiny?

It can and does withstand scrutiny. What are you trying to say here?

HIS theory is pure "philosophy", and based on Junk Science.

ROFL!!! Okay, you keep believing that, while the rest of the world keeps doing research, which to date has overwhelmingly confirmed evolution.

But just for giggles, feel free to give some specific examples, in detail, of the "junk science" you mistakenly believe evolutionary biology is "based on"...

Let's test whether you really have any idea what you're talking about, or whether you've made the mistake of reading creationist sources -- and believing them. Note: Trying to "learn" about science from creationist sources is like trying to "learn" about conservatism by watching Michael Moore movies.

[Doctor Stochastic]

So, was it two or seven of each of these on the ship?

[MHalblaub]

Something for all to think about before you defend your theories to much.

...but those who uphold it dogmatically [a system] . . . are adopting the very reverse of that critical attitude which in my view is the proper one for the scientist. In point of fact, no conclusive disproof of a theory can ever be produced; . . . If you insist on strict proof (or strict disproof) in the empirical sciences, you will never benefit from experience, and never learn from it how wrong you are.


Karl R. Popper, The Logic of Scientific Discovery
"Evolution", "ID" or "Creationism" is never mentioned.

[restornu]

Fact, Fable, and Darwin, Part 2

,

[Youngblood]

A happy birthday to Messrs. Darwin and Lincoln, two giants of the nineteenth or any other century!

[phoenix0468]

hey Peter, I think there are quite a few here who are both Darwinists and Christian. In fact most who would respond to a thread like this would fall under that category. I on the other hand am the anomaly. I am and agnostic anti-Darwinist. My reasons are just what is presented in this article and in a subsequent article posted on 2-11. Darwin based most of his theory on conjecture and circumstantial evidence rather than hard facts. I do not dispute many of the facts of evolutionary biology regarding things such as adaptations and mutations withing species, but the clear and evident boundaries between species makes outright and spontaneous evolution seem far fetched. Without some outside influence, it just does not seem possible to me that a fish came from a bacteria or a human from a fish. And this is probably the most basic of "facts" of evolutionary biology. This "fact" also happens to be the one that is most weakly supported by evidence. Thus we have the theory of the "missing links" of many species' evolution. So before you go attacking the forum as a whole as you have done, sit back and read the posts presented. Then you can post your rant. Otherwise, you may just end up looking foolish.

[RadioAstronomer]

I proved him wrong today like 3 other times on three other things.

Huh? Here is what is meant when the term theory is used in science:

http://en.wikipedia.org/wiki/Theory

"In common usage a theory is often viewed as little more than a guess or a hypothesis. But in science and generally in academic usage, a theory is much more than that. A theory is an established paradigm that explains all or many of the data we have and offers valid predictions that can be tested. In science, a theory can never be proven true, because we can never assume we know all there is to know. Instead, theories remain standing until they are disproven, at which point they are thrown out altogether or modified slightly.

Theories start out with empirical observations such as “sometimes water turns into ice.” At some point, there is a need or curiosity to find out why this is, which leads to a theoretical/scientific phase. In scientific theories, this then leads to research, in combination with auxiliary and other hypotheses (see scientific method), which may then eventually lead to a theory. Some scientific theories (such as the theory of gravity) are so widely accepted that they are often seen as laws. This, however, rests on a mistaken assumption of what theories and laws are. Theories and laws are not rungs in a ladder of truth, but different sets of data. A law is a general statement based on observations."

For Laws:

"A well-known example is that of Newton's law of gravity: while it describes the world accurately for most pertinent observations, such as of the movements of astronomical objects in the solar system, it was found to be inaccurate when applied to extremely large masses or velocities. Einstein's theory of general relativity, however, accurately handles gravitational interactions at those extreme conditions, in addition to the range covered by Newton's law. Newton's formula for gravity is still used in most circumstances, as an easier-to-calculate approximation of gravitational law. A similar relationship exists between Maxwell's equations and the theory of quantum electrodynamics; there are several such cases. This suggests the (unanswered) question of whether there are any ultimately true physical laws, or whether they are all just cases where our sensory and rational apparatus have generated mathematically simple approximations, valid within the range of normal human experience, to unobtainable true formulas."

Let me post my example of gravity:

A little history here: Newton’s Law of Universal Gravitation

“Every object in the universe attracts every other object with a force directed along the line of centers for the two objects that is proportional to the product of their masses and inversely proportional to the square of the separation between the two objects.”

F=Gm₁m₂/r²

Where:

F equals the gravitational force between two objects
m₁ equals the mass of the first object
m₂ equals the mass of the second object
R equals the distance between the objects
G equals the universal constant of gravitation = (6.6726 )* 10^-11 N*m²/kg² (which is still being refined and tested today)

(BTW this is a simple form of the equation and is only applied to point sources. Usually it is expressed as a vector equation)

Even though it works well for most practical purposes, this formulation has problems.

A few of the problems are:

It shows the change is gravitational force is transmitted instantaneously (Violates C), assumes an absolute space and time (this contradicts Special Relativity), etc.

Enter Einstein’s General Theory of Relativity

In 1915 Einstein developed a new theory of gravity called General Relativity.

A number of experiments showed this theory explained some of the problems with the classical Newtonian model. However, this theory like all others is still being explored and tested.

And finally:

From an NSF abstract:

“As with all scientific knowledge, a theory can be refined or even replaced by an alternative theory in light of new and compelling evidence. The geocentric theory that the sun revolves around the earth was replaced by the heliocentric theory of the earth's rotation on its axis and revolution around the sun. However, ideas are not referred to as "theories" in science unless they are supported by bodies of evidence that make their subsequent abandonment very unlikely. When a theory is supported by as much evidence as evolution, it is held with a very high degree of confidence.

In science, the word "hypothesis" conveys the tentativeness inherent in the common use of the word "theory.' A hypothesis is a testable statement about the natural world. Through experiment and observation, hypotheses can be supported or rejected. At the earliest level of understanding, hypotheses can be used to construct more complex inferences and explanations. Like "theory," the word "fact" has a different meaning in science than it does in common usage. A scientific fact is an observation that has been confirmed over and over. However, observations are gathered by our senses, which can never be trusted entirely. Observations also can change with better technologies or with better ways of looking at data. For example, it was held as a scientific fact for many years that human cells have 24 pairs of chromosomes, until improved techniques of microscopy revealed that they actually have 23. Ironically, facts in science often are more susceptible to change than theories, which is one reason why the word "fact" is not much used in science.

Finally, "laws" in science are typically descriptions of how the physical world behaves under certain circumstances. For example, the laws of motion describe how objects move when subjected to certain forces. These laws can be very useful in supporting hypotheses and theories, but like all elements of science they can be altered with new information and observations.

Those who oppose the teaching of evolution often say that evolution should be taught as a "theory, not as a fact." This statement confuses the common use of these words with the scientific use. In science, theories do not turn into facts through the accumulation of evidence. Rather, theories are the end points of science. They are understandings that develop from extensive observation, experimentation, and creative reflection. They incorporate a large body of scientific facts, laws, tested hypotheses, and logical inferences. In this sense, evolution is one of the strongest and most useful scientific theories we have."

[RadioAstronomer]

So, what's going on here? He still can't explain HOW life came about. If Ernst Mayr is the "end all" of neo-Darwinism, how can his version of Darwin's theory be so weak as to not withstand basic scrutiny? HIS theory is pure "philosophy", and based on Junk Science.

Whoops! WRONG!! Evolution does not address origins. That is a different theory.

[RadioAstronomer]

I think it's worth a ping because of the credibility the AEI has among conservatives. This is no fringe organization, but a highly influential thinktank. We need the regulars here to refute this nonsense before it becomes respectable.

Agree completely.

[RadioAstronomer]

Just don't kid yourself you're learning any science.

Well said.

[Doctor Stochastic]

Rather than Creationist nonsense becoming respectable, it's more likely that the AEI (and other Conservative institutions) will be seen as anti-science. When issues needing scientific input come up politically (ABM, greenhouse, public health, etc.), Conservatives will not be seen as credible participants. Things have progressed much farther than most people think.

[LauraleeBraswell]

In this sense, evolution is one of the strongest and most useful scientific theories we have."

OPINION

How did an EYE evolve then?

and actually, there are alternatives to evolution. No, not creationism.

I'm an adherant to intelligent design. Phi- the golden ratio.