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From the Captain and Tennille website, here is the explanation for how their version of "Muskrat Love" came about:

The third single released from Song of Joy was an unlikely number called "Muskrat Love." Loved and loathed, the revamped version of country composer/singer Willis Alan Ramsey’s "Muskrat Candlelight" was another million seller for the duo. "We were driving around one night when we heard (the group) America’s version of the song on the radio," remarks Toni. "I said to Daryl ‘did you hear that? I swear they’re singing about muskrats.’ I had to know what the lyrics were, so the next day we went out and found the sheet music" Toni reminisces. "I said to Daryl, this song is hysterical, why don’t we add it to our club-act? And people went nuts for it." Daryl continues "We just threw it on the second album because we had room for one more tune. We knew it was a hit in the clubs, but we had no idea it would be a hit record." The label had no intention of releasing a third single from the album, but after a radio station in Madison, Wisconsin called the A&R people at A&M begging them for a copy, telling them the phones were ringing off the hooks for it, a 45 was issued and became their third biggest hit.