The various Ensatina salamanders of the Pacific coast all descended from a common ancestral population. As the species spread southward from Oregon and Washington, subpopulations adapted to their local environments on either side of the San Joaquin Valley. From one population to the next, in a circular pattern, these salamanders are still able to interbreed successfully. However, where the circle closes -- in the black zone on the map in Southern California -- the salamanders no longer interbreed successfully. The variation within a single species has produced differences as large as those between two separate species