There are also pass along copies measured by independent firms. The total readers of a large newspaper like USA Today is about 5 million or so readers.
Newspapers know what their demographics are. As you are aware, advertisers only use advertising vehicles that match their needs. BMW does not advertise in a newspaper in which the readers have little discretionary income.
A color ad in USA Today is much more expensive than any popdown ad in cyberspace.....tons more expensive because the advertiser values the newspaper reader much much more than the cyberreader for many many reasons.