Great point. Millions of bison over tens of thousands of years sure are going to do a great job of fertilizing the soil.
It's not just poop. It's grazing grasses to facilitate forbs with their microbial relationships. Grazing also forces cyclical root die-back which adds organic matter.
Animal disturbance is not only elemental to soil formation, it inhibits successional establishment of perennial cover to retain the region in grasslands. That too builds a deeper soil.
Over 70% of agricultural soils are former grasslands.