They also try to use a coupon for both the initial purchase, and for anything bought with the Register Rewards.
Presumably, there also are still a few manufacturer's mail-in rebates around, too.
Here's official information from the Walgreens website:
As a customer-focused retailer, Walgreens encourages the use of coupons by our customers in our retail stores, in accordance ...
http://www.walgreens.com/topic/help/generalhelp/coupon_policy_main.jsp
Register Rewards | Walgreens
The Walgreens Jingle Ca$h coupon value cannot exceed the total purchase amount.
No cash back and no cash value for the Walgreens Jingle Ca$h coupon. ...
http://www.walgreens.com/topic/save-money/register-rewards.jsp