This seems to be saying that population density, probably clustering, was higher in the hunter-gatherer north.
I thought that interesting as, when we get to recorded times, there is contention as to how populous were the groups we call Germanic and Mongolian. IOW, did they come to prevail much out of force of numbers.
It does make a bit of sense, if I say so myself, as shepherding societies tend to be patriarchal and worship male gods (including the Hebrew God) and agricultural societies tend toward matriarchies and Earth mother flim-flam.