They also used MUSEUM SAMPLES (of questionable provinence I might add).
Article we are discussing is quite recent and is based on thousands of samples taken recently.
I think the old conclusions have been overturned.
BTW, the discovery that the San people are apparantly the source of the most ancient lineages for Europeans and Chinese (and everybody else outside of Africa) is NEW STUFF.
Nils Vander Post suggested as much 3/4 of a century ago.
Genetic studies often seem to me to conflict with each other.
As to San being ancestral to us honkies, see my post about the Venus of Willendorf.