There are actually two complimentary papers being discussed. Here is a related quote about the second one:
"In the second study, Terry Root of Stanford University, California, and colleagues also report a temperature-related fingerprint in the behaviour of a range of species."
"They found the changes were most marked at high latitudes and high altitudes, where the largest temperature changes are predicted."
So the average is less (perhaps, given that I haven't read the actual papers) than in the areas most affected by warming. And who's supposed to be very surprised by that?
Here's another perspective:
Warming planet shifts life north and early
""Animals and plants, of all different types, around the globe are feeling the effects of global warming of only 0.6 ºC over the last 100 years," says Root. She says that although wildlife has adapted to climate change in the past, the current rate is ten times faster than previous global swings in temperature."