[tallhappy]

...histones had to evolve stepwise into this one and only function, all while being completely nonfunctional until they reached that magic end result - homologs such as the protamines ...

Protamines are not histone homologs.

[Senator Bedfellow]

"Such as" is probably a bad choice of phrasing - were I to rewrite, it would be "akin to" or "similar to".