What is the working definition of Culture here?
Depends, doesn't it?
If you are a person living within an extant culture, it is the aquarium in which you swim, the commonly held philosophy, viewpoint and/or belief system which influences all of your perceptions.
If you want the definition of culture from the historical viewpoint, culture is that which remains once all of them who lived it, are gone.
If I were to evaluate a new hire in terms of culture, I would want to know how well that person was adapting to the culture in their potential work environment.