...and where is your proof that the beginnings of life had a fitness cost profile much different from the current electronic models?
The switch, as usual. We are not talking about fitness cost in the beginning of life, and the article does not deal with the beginning of life either. We are discussing evolution, the time after the beginning of life. The problem with the beginning of life is not fitness cost, it is the absolute impossibility of arranging some millions of molecules into at least half a million DNA pairs in the exact way necessary to produce a living thing.
As to fitness cost, I already gave it to you in the post you are responding to " However, in real life there is a fitness cost of non-useful organs, DNA, etc. It takes energy, food, etc. to keep such useless things alive so there is definitely a fitness cost." Further you already agreed that even a small fitness differential will result in the more fit organisms overcoming the less fit.