But that mutation rate is directed at specific areas at specific rates. That is not Darwinian. You missed this in Shapiro's article.
Darwin himself acknowledged this point in later editions of Origin of Species, where he wrote about natural "sports" or "...variations which seem to us in our ignorance to arise spontaneously. It appears that I formerly underrated the frequency and value of these latter forms of variation, as leading to permanent modifications of structure independently of natural selection." (6th edition, Chapter XV, p. 395).