Why keep proving that you cannot read. This paper talked about known deleterious mutations in one species being subsequently discovered in another species in the wild. The conclusion was ...Thus, compensating mutations must occur and become fixed very frequently in populations.
The authors concluded that some other presently unknown mutation(s) had to compensate for the presently known deleterious mutation in the wild species. The harmful mutations were fixed in the genome of the wild species.
I think your analysis of the Science article is badly flawed. Nice obfuscation, though.