The wording of the article indicates that the "not for on-road use" is a condition separate and distinct from the "no title". As in "these are not road-legal, period".
In case you were planning to rumble up to church on Christmas morning and scare the little old ladies in the parking lot in one of these things, they will not be considered roadworthy.
Which makes me wonder about the National Guard Hummers that we all see on the highway as they go to and from their weekend training.
I know the mil trucks don’t have to meet some of the MVSS, NHTSA, and EPA standards for regular on-highway motor vehicles. But if your state licensing is good with that, enjoy the ride.