About a year ago I attended a fascinating lecture at the Carnegie Institution in DC. The presenter spoke of the shrinking X chromosome, and then described a fascinating level of complexity with another species that, if I remember correctly, had several X, one X or no X but behaved as a male. It was very puzzling. I think it might have been naked mole rats, but I have to go to work soon, so maybe you would want to Google that.
He was in the pool!